The tertiary consumers in a forest build 6 kJ/m2 of new tissue a year. How much must the producers make, at 10% efficiency per step? How does the answer change at the measured average of 5.92%?
1
Count the steps
Tertiary consumers are level 4, three steps above the producers. Write E for the producers’ production in kJ/m2 a year. Each step keeps 0.1 of the level below, so three steps keep 0.13 of E.
E(0.1)3=6
2
Evaluate the power
0.13=0.001, so the tertiary level keeps one thousandth of the producers’ production.
0.001E=6
3
Divide
Divide both sides by 0.001 to leave E alone.
E=0.0016
4
Evaluate
The producers must make 6,000 kJ/m2 a year.
E=6000
5
Use the measured average
At 5.92% per step the producers would need 6/0.05923, nearly 29,000 kJ/m2 a year. A drop in efficiency from 10% to about 6% means almost five times as much plant production for the same predators.
0.059236≈28919
6
Check by going up
Going up from 6,000 at 10% a step gives 600,60 and 6, as required. The trophic pyramid in Biology shows those four levels and a 10% transfer at each step.
Result
6,000 kJ/m2 a year at 10% per step. At the measured 5.92% the producers would need nearly 29,000 kJ/m2 a year, almost five times as much.
Your turn
1. One like the example
Secondary consumers in a grassland build 50 kJ/m2 a year. At 10% per step, how much must the producers make?
Show the answer and explanation
5,000 kJ/m2 a year.
Two steps, so E(0.1)2=50,0.01E=50 and E=50/0.01=5,000.
E(0.1)2=500.01E=50E=0.0150E=5000
2. A variation
A coastal sea makes 120 kJ/m2 of phytoplankton a year, and each step passes on 8%. How much reaches the fourth trophic level?
Show the answer and explanation
About 0.0614 kJ/m2 a year.
Three steps up, so 120×0.083=120×0.000512≈0.0614. Multiplying by 0.08×3 instead of 0.083 is the slip to avoid.
120(0.08)3≈0.0614
3. Somewhere new
People can eat grain directly or eat cattle fed on grain. At 10% per step, how much more grain does the second diet need for the same energy, and how much more would a diet two steps up need?
Show the answer and explanation
Ten times as much one step up, and a hundred times two steps up.
Eating one step higher divides the energy by the efficiency, 1/0.1=10, and two steps divide it by 0.12,giving 100. The 10% is the rule of thumb, so the true factor depends on the animal. Mammals such as cattle keep a warm body and spend much of what they absorb on body heat.
0.11=100.121=100
Keep exploring
The trophic pyramid tool opens with producers at 6,000 kJ/m2 a year and 600,60 and 6 above them, each transfer 10%.